Regeane is Maeniel's mate (he's the long-lived werewolf leader of the pack, whose earlier life was featured in Night of the Wolf). Once thawed, Regeane confronts a demented abbot and a gang of (literal) cutthroats to save him. The werewolves and the Saxon head for Geneva to pledge allegiance to Charlemagne, who's about to cross the Alps to challenge King Desederius of the Lombards for control of northern and central Italy. Soon Maeniel is in Desederius's territory and in danger. Regeane follows, despite his prohibition. They're fated to reencounter Regeane's sniveling cousin Hugo, who seeks revenge. He has become host to a powerful bear spirit who wants the wolves for his own purposes. The new Hugo has a lot in common with the Steve Martin/Lily Tomlin character in All of Me; he provides comic leavening to the sometimes grim action. Other returning characters include Pope Hadrian's tough, practical, but vulnerable mistress Lucilla; her protégé, the singer Dulcinia; and the ageless werewolf earth-mother Matrona.
